Leioclema Ulrich, 1882

Type species: Callopora punctata Hall, 1858

The alternate spelling Lioclema is widely used for this genus.

Synonyms Author Type Species
Leioclemina Vinassa de Regny, 1921 Lioclema ramulosum Bassler, 1906
Thallostigma Hall, 1883 Thallostigma intercellata Hall, 1883

This genus includes the species:

  1. Leioclema abacanensis
  2. Schoenmann, 1926 Devonian Givetian? Russia
  3. Leioclema abnorme
  4. (Morozova, 1969) Triassic Carnian Russia
  5. Leioclema acanthinum
  6. Bigey, 1986 Silurian Pridoli France
  7. Leioclema aciculata
  8. Modzalevskaya, 1980 Silurian Russia Tuva
  9. Leioclema aculeata
  10. Nekhoroshev, 1977 Devonian Givetian Russia?
  11. Leioclema adsuetum
  12. Ernst, Brett & Wilson, 2019 Silurian Llandovery Aeronian USA New York
  13. Leioclema aequilaterale
  14. Kopajevich, 1984 Devonian Eifelian Mongolia
  15. Leioclema? afghana
  16. Termier & Termier, 1971 Lower Carboniferous Afghanistan
  17. Leioclema alpenense
  18. Duncan, 1939 Devonian Givetian USA
  19. Leioclema altaica
  20. Yaroshinskaya, 1997 Silurian Russia Gornyi Altai
  21. Leioclema amplexum
  22. Kopajevich, 1984 Silurian Ludlovian-Pridolian Mongolia
  23. Leioclema angulata
  24. Yaroshinskaya, 1990 Silurian Russia
  25. Leioclema araneum
  26. Ulrich, 1890 Lower Carboniferous Visean-Namurian USA Illinois, Kentucky
  27. Leioclema avonense
  28. Lee, 1912 Lower Carboniferous Tournaisian England
  29. Leioclema blandum
  30. Jaroshinskaya, 1970 Early Devonian Mongolia
  31. Leioclema breve
  32. Liu, 1980 Middle Devonian China
  33. Leioclema bugusunicum
  34. Nekhoroshev, 1948 Middle Devonian Russia
  35. Leioclema caelata
  36. Liu, 1976 Lower Permian China
  37. Leioclema catenatus
  38. Yang & Xia, 1976 Silurian Pridoli China Yunnan
  39. Leioclema celebratum
  40. (Morozova, 1961) Devonian Givetian-Frasnian Russia, Mongolia, Poland
  41. Leioclema cellulosum
  42. (Hall, 1879) Early Devonian Lochkovian USA Tennessee
  43. Leioclema? choshiensis
  44. Sakagami, 1965 Permian Guadalupian Japan
  45. Leioclema clarum
    (Homonym?)
  46. Liu, 1993? Devonian China Sichuan
  47. Leioclema claudestinum
  48. Liu, 1993? Devonian China Sichuan
  49. Leioclema confertiporum
  50. (Hall, 1883) Middle Devonian Givetian USA New York
  51. Leioclema crassimurale
  52. (Romanchuk, 1979) Early Devonian Russia Amur
  53. Leioclema crassiparietum
  54. Ernst & Königshof, 2010 Middle Devonian Givetian Western Sahara
  55. Leioclema crenulata
  56. Ernst, 2008 Middle Devonian Germany
  57. Leioclema cruciatum
  58. Yang & Lu, 1962 Lower Permian China
  59. Leioclema crustulum
  60. (Astrova, 1965) Silurian Ludlow Ukraine Podolia
  61. Leioclema cystiramoides
  62. Kopajevich, 1984 Early Devonian Mongolia
  63. Leioclema daminshanense
  64. Liu, 1980 Late Devonian China
  65. Leioclema decipiens
  66. (Hall, 1883) Middle Devonian Givetian USA New York
  67. Leioclema densiporum
  68. Owen, 1965 Silurian Llandovery England
  69. Leioclema densum
  70. (Hall, 1881) Middle Devonian Givetian USA New York
  71. Leioclema digitatum
  72. (Hall, 1881) Middle Devonian Givetian USA New York
  73. Leioclema? distans
  74. (Whidborne, 1896) Devonian England
  75. Leioclema dozierense
  76. (Moore, 1940) Permian Guadalupian USA Texas
  77. Leioclema echinatum
  78. (Trizna, 1958) Carboniferous Mississippian Visean Russia Kuznetsk Basin
  79. Leioclema editum
  80. (Morozova, 1961) Late Devonian Frasnian Russia
  81. Leioclema elasmaticum
  82. Boardman, 1960 Early-Middle Devonian USA New York, Oklahoma
  83. Leioclema elegans
  84. Ernst, 2008 Early Devonian Pragian Czech Republic
  85. Leioclema emarginatum
  86. (Kopajevich, 1984) Silurian Ludlow-Pridoli Mongolia
  87. Leioclema eoexplanatum
  88. Modzalevskaya, 1979 Silurian Llandovery Russia
  89. Leioclema exileramosa
  90. Nekhoroshev, 1977 Devonian Givetian Russia
  91. Leioclema exilis
  92. (Pushkin, 1976) Silurian Pridolian Belarus
  93. Leioclema explanodeum
  94. Modzalevskaya, 1979 Silurian Ludlow Russia
  95. Leioclema exsul
  96. (Hall, 1876) Silurian Wenlock USA Indiana
  97. Leioclema florea
  98. (Modzalevskaya, 1965) Middle Devonian Russia
  99. Leioclema foliatum
  100. Ulrich, 1889 Carboniferous Mississippian Visean USA Illinois
  101. Leioclema ganiense
  102. (Astrova, 1968) Early Devonian Emsian Russia
  103. Leioclema globosum
  104. (Astrova, 1954) Early Devonian Lochkovian Russia?
  105. Leioclema globulare
  106. (Bassler, 1906) Silurian Wenlock USA New York
  107. Leioclema glomeratum
  108. (Astrova, 1965) Silurian Ludlow Russia
  109. Leioclema gracilenta
  110. Liu, 1980 Middle Devonian China
  111. Leioclema gracillimum
  112. Ulrich, 1890 Carboniferous Mississippian Tournaisian-Visean USA Illinois, Iowa, Ohio
  113. Leioclema guanniaoheense
  114. Liu, 1980 Middle Devonian China
  115. Leioclema halloporoides
  116. Owen, 1962 Silurian Ludlow England
  117. Leioclema heckeri
  118. (Trizna, 1948) Permian Cisuralian Artinskian Russia Urals
  119. Leioclema heitaiense
  120. (Yang, 1956) Devonian Givetian-Frasnian China, Russia
  121. Leioclema hirsutum
  122. Moore, 1929 Carboniferous Pennsylvanian Gzhelian USA Texas
  123. Leioclema implicatum
  124. (Kopajevich, 1984) Silurian Ludlow-Pridoli Mongolia
  125. Leioclema incompositum
  126. (Duncan, 1939) Middle Devonian Givetian USA Michigan
  127. Leioclema incrustans
  128. (Nekhoroshev, 1948) Devonian Givetian-Frasnian Russia
  129. Leioclema indentatum
  130. Wyse Jackson, 1996 Carboniferous Mississippian Visean Ireland
  131. Leioclema intercellatum
  132. (Hall, 1881) Middle Devonian Eifelian USA Indiana
  133. Leioclema invalida
  134. Liu, 1987 Late Ordovician China
  135. Leioclema? involvens
  136. (Hall & Simpson, 1887) Middle Devonian Givetian USA New York
  137. Leioclema irinae
  138. (Astrova, 1959) Silurian Wenlock Russia Tuva
  139. Leioclema irregulatum
  140. (Yang, 1956) Middle Devonian Givetian China
  141. Leioclema ivanovae
  142. (Morozova, 1961) Late Devonian Famennian Russia
  143. Leioclema jeni
  144. (Yang, 1956) Devonian Givetian-Frasnian Mongolia
  145. Leioclema jinshuiense
  146. Liu, 1980 Early Devonian China
  147. Leioclema khalfini
  148. Jaroshinskaya, 1970 Early Devonian Russia? Altai
  149. Leioclema kisiltschinense
  150. (Volkova, 1969) Devonian Russia
  151. Leioclema kitajense
  152. (Astrova, 1965) Silurian Llandovery-Wenlock Ukraine Podolia
  153. Leioclema kobayashii
  154. Sakagami, 1962 Carboniferous Mississippian Tournaisian Japan
  155. Leioclema kusmense
  156. Tolokonnikova, 2012 Late Devonian Famennian Russia Kuznetsk Basin
  157. Leioclema leioclemaformis
  158. (Yang, & Xia, 1975) Late Triassic China
  159. Leioclema liuchingense
  160. (Hu, 1965) Middle Devonian China
  161. Leioclema maculatum
  162. (Nekhoroshev, 1948) Middle Devonian Givetian Russia
  163. Leioclema manchuriense
  164. (Yang, 1956) Middle Devonian Givetian China
  165. Leioclema maslovi
  166. (Astrova, 1959) Silurian Wenlock Russia Tuva
  167. Leioclema mcgerriglei
  168. (Fritz, 1940) Early Devonian Pragian Canada
  169. Leioclema meisteri
  170. (Troizkaya, 1979) Late Devonian Famennian Kazakhstan
  171. Leioclema microporum
  172. (Hall, 1881) Middle Devonian Givetian USA New York
  173. Leioclema microporum
    (Homonym)
  174. Sakagami, 1964 Permian Cisuralian Japan
  175. Leioclema minussinense
  176. (Morozova, 1961) Late Devonian Frasnian Russia
  177. Leioclema minutissimum
  178. (Nicholson, 1875) Middle Devonian Givetian Canada
  179. Leioclema? minutum
  180. (Rominger, 1866) Late Devonian Frasnian USA Iowa
  181. Leioclema mirum
  182. (Morozova, 1961) Middle Devonian Givetian Russia Kuznetsk Basin
  183. Leioclema modzalevskajae
  184. Yaroshinskaya, 1990 Silurian Russia
  185. Leioclema mongraiensis
  186. Sakagami, 1968 Permian Cisuralian Artinskian Thailand
  187. Leioclema morozovae
  188. Yaroshinskaya, 1990 Silurian Russia
  189. Leioclema multaculeatum
  190. (Hall, 1884) Middle Devonian Givetian USA New York
  191. Leioclema multiacanthoporum
  192. (Astrova, 1968) Early Devonian Emsian Russia, Mongolia
  193. Leioclema multum
  194. (Volkova, 1969) Middle Devonian Givetian Russia Altai
  195. Leioclema multum
    (Homonym)
  196. (Yang & Xia, 1975) Late Triassic Carnian China
  197. Leioclema muratae
  198. Sakagami, 1964 Permian Guadalupian Japan
  199. Leioclema neoavonense
  200. Yang & Lu, 1962 Permian Cisuralian China
  201. Leioclema netshlavense
  202. (Astrova, 1964) Early Devonian Russia, Mongolia
  203. Leioclema nivale
  204. (Reed, 1922) Late Devonian Frasnian India
  205. Leioclema nonakae
  206. Sakagami, 1963 Permian Cisuralian Artinskian-Kungurian Japan
  207. Leioclema numerosum
  208. (Morozova, 1961) Late Devonian Famennian Russia
  209. Leioclema occidens
  210. (Hall & Whitfield, 1873) Late Devonian Frasnian USA Iowa
  211. Leioclema orbicularis
  212. Modzalevskaya, 1953 Ordovician Llandeilo-Caradoc Russia
  213. Leioclema parasiticum
  214. (Hall, 1879) Early Devonian Lochkovian USA New York
  215. Leioclema parvum
  216. (Astrova, 1964) Early Devonian Lochkovian Russia
  217. Leioclema passitabulatum
  218. (Duncan, 1939) Middle Devonian Givetian USA Michigan
  219. Leioclema pauciporum
  220. Yang & Xia, 1976 Middle Devonian China
  221. Leioclema perexiguum
  222. (Astrova, 1959) Silurian Wenlock Russia Tuva
  223. Leioclema pilulum
  224. (Yang & Xia, 1975) Late Triassic China
  225. Leioclema plicatile
  226. (Volkova, 1969) Late Devonian Frasnian Russia? Altai
  227. Leioclema polenovi
  228. (Nekhoroshev, 1948) Middle Devonian Givetian Russia
  229. Leioclema ponderosum
  230. (Hall, 1874) Early Devonian Lochkovian USA New York
  231. Leioclema porosum
  232. Crockford, 1947 Carboniferous Mississippian Visean Australia Queensland
  233. Leioclema praepassitabulatum
  234. Yaroshinskaya, 1990 Silurian Russia
  235. Leioclema pressulum
  236. (Troizskaya, 1968) Late Devonian Frasnian Russia
  237. Leioclema proprium
  238. Jaroshinskaya, 1970 Early Devonian Russia? Altai
  239. Leioclema pulchellum
  240. (Ulrich, 1893) Ordovician Caradoc USA Minnesota
  241. Leioclema pulchellum
    (Homonym)
  242. (Ulrich & Bassler, 1913) Silurian-Devonian USA
  243. Leioclema pulchelliforme
  244. Pushkin, 1976 Silurian Ludlow Belarus
  245. Leioclema punctatum
  246. (Hall, 1858) Mississippian USA (Iowa)
  247. Leioclema punctillatum
  248. (Winchell, 1866) Middle Devonian Givetian USA Michigan
  249. Leioclema pushmatahense
  250. (Harlton, 1933) Carboniferous Pennsylvanian Bashkirian USA Oklahoma
  251. Leioclema qujingense
  252. Hu & Fan, 1985 Silurian Pridoli China Yunnan
  253. Leioclema ramosum
  254. (Nekhoroshev, 1948) Late Devonian Famennian Russia
  255. Leioclema ramulosum
  256. (Bassler, 1906) Silurian Llandovery-Wenlock USA New York
  257. Leioclema rectimurale
  258. Bigey, 1986 Silurian Pridoli France
  259. Leioclema? reeftonensis
  260. (Allan, 1935) Early Devonian Pragian New Zealand
  261. Leioclema regularis
  262. Zheng, 1990 Ordovician China
  263. Leioclema salairense
  264. (Morozova, 1961) Middle Devonian Eifelian Russia
  265. Leioclema segregatum
  266. (Hall, 1881) Middle Devonian Givetian USA New York
  267. Leioclema shumardi
  268. Girty, 1908 Permian Guadalupian Capitanian USA Texas
  269. Leioclema slygapensis
  270. (Fritz, 1944) Late Devonian Frasnian USA New Mexico
  271. Leioclema speciosum
  272. Hu, 1982 Silurian Wenlock China
  273. Leioclema subasperum
  274. Modzalevskaya, 1980 Silurian Ludlow Russia Tuva
  275. Leioclema subglobosum
  276. Ulrich, 1890 Carboniferous Mississippian Tournaisian USA Iowa
  277. Leioclema subramosa
  278. (Ulrich & Bassler, 1913) Silurian-Devonian USA, Russia
  279. Leioclema subtile
  280. (Hall, 1881) Middle Devonian Givetian USA New York
  281. Leioclema tapsaense
  282. (Astrova, 1959) Silurian Wenlock Russia Tuva
  283. Leioclema tenuiramum
  284. (Bassler, 1923) Silurian Pridoli USA West Virginia
  285. Leioclema textile
  286. (Trizna, 1958) Carboniferous Mississippian Tournaisian Russia Kuznetsk Basin
  287. Leioclema trematoporoides
  288. (Astrova, 1954) Early Devonian Lochkovian Russia
  289. Leioclema triviale
  290. Liu, 1980 Late Devonian China
  291. Leioclema tschuyaense
  292. (Volkova, 1969) Devonian Russia
  293. Leioclema tubulosum
  294. (Nekhoroshev, 1956) Carboniferous Mississippian Tournaisian Russia? Altai
  295. Leioclema tulungense
  296. (Yang & Xia, 1975) Late Triassic Carnian China
  297. Leioclema tungi
  298. (Yang, 1956) Middle Devonian Givetian China
  299. Leioclema turanense
  300. (Modzalevskaya, 1968) Silurian Ludlow Russia Tuva
  301. Leioclema tuvaense
  302. (Astrova, 1959) Silurian Wenlock Russia Tuva
  303. Leioclema? uzuraense
  304. Sakagami, 1964 Carboniferous Mississippian Visean Japan
  305. Leioclema varium
  306. (Astrova, 1959) Silurian Wenlock-Ludlow Russia Tuva
  307. Leioclema vassinense
  308. (Morozova, 1961) Late Devonian Famennian Russia
  309. Leioclema wachsmuthi
  310. Ulrich, 1890 Carboniferous Mississippian Tournaisian USA Iowa
  311. Leioclema waschurovae
  312. Nekhorosheva, 1988 Silurian Ludlow Tajikistan
  313. Leioclema yakovlevi
  314. (Schoenmann, 1926) Devonian Russia, Mongolia
  315. Leioclema yakovlevi palmipedalis
  316. Schoenmann, 1926 Middle Devonian Givetian Russia
  317. Leioclema yakovlevi tenuimurinum
  318. (Volkova, 1969) Devonian Russia
  319. Leioclema yakovlevi vulgatum
  320. (Volkova, 1969) Devonian Russia
  321. Leioclema yuzhenshanense
  322. Liu, 1980 Early-Middle Devonian China
  323. Leioclema zonale
  324. (Kopajevich, 1984) Devonian Givetian-Frasnian China
 

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Edited by Phil Bock
Modified on 13/02/2016
Modified on 13/8/2017 (several species added or removed)
Modified on 12/2/2019 (Added L. adsuetum)
This URL is http://bryozoa.net/trepostomatida/leioclemidae/leioclema.html