Heterotrypa Nicholson, 1879

Type species: Monticulipora frondosa d'Orbigny, 1850

Synonyms Author Type Species
Dekayella Ulrich, 1882 Dekayella obscura Ulrich, 1883 (designation invalid?)

This genus includes the species:

  1. Heterotrypa aculeata
  2. Hu, 1986 Late Ordovician Ashgill China
  3. Heterotrypa affinis
  4. (Ulrich, 1890) Late Ordovician Ashgill USA (Illinois)
  5. Heterotrypa astrovae
  6. Pushkin, 1976 Late Ordovician Caradoc Belarus
  7. Heterotrypa compressa
  8. Brood, 1978 Late Ordovician Ashgill Sweden Gotland
  9. Heterotrypa crassimuralis
  10. (Astrova, 1965) Middle Ordovician Arctic Russia
  11. Heterotrypa cystata
  12. (Cumings, 1901) Late Ordovician Ashgill USA (Indiana)
  13. Heterotrypa? dalpiazzi
  14. Vinassa de Regny, 1921 Late Ordovician Italy
  15. Heterotrypa definita
  16. Dyer, 1925 Late Ordovician Ashgill Canada (Ontario)
  17. Heterotrypa enormis
  18. Astrova, 1959 Silurian Ludlovian Russia
  19. Heterotrypa exovaria
  20. Marintsch, 1998 Middle Ordovician USA (Tennessee)
  21. Heterotrypa extensa
  22. Fan & Hu, 1985 Late Silurian China
  23. Heterotrypa foliacea
  24. (Ulrich & Bassler, 1904) Late Ordovician Caradoc USA (Kentucky)
  25. Heterotrypa frondosa
  26. (d'Orbigny, 1850) Late Ordovician Caradoc USA (Ohio, Indiana, Kentucky)
  27. Heterotrypa georgii
  28. Modzalevskaya, 1968 Silurian Ludlovian Russia
  29. Heterotrypa? girvanensis
  30. (Nicholson, 1881) Late Ordovician Caradoc Scotland
  31. Heterotrypa humensis
  32. Ross, 1961 Silurian Australia (NSW)
  33. Heterotrypa incelebrata
  34. Modzalevskaya, 1968 Silurian Ludlovian Russia
  35. Heterotrypa incrustans
  36. Pushkin, 1976 Silurian, Llandovery Belarus
  37. Heterotrypa lata
  38. Kopajevich, 1984 Late Ordovician Ashgill Mongolia
  39. Heterotrypa magnopora
  40. Boulange, 1964 Late Ordovician Ashgill France
  41. Heterotrypa magnopora
    (Homonym)
  42. Marintsch, 1998 Middle Ordovician USA (Tennessee)
  43. Heterotrypa meafordensis
  44. Fritz, 1975 Late Ordovician Ashgill Canada
  45. Heterotrypa? megacanthopora
  46. (Owen, 1960) Silurian Ludlovian Wales
  47. Heterotrypa microstigma
  48. Cumings & Galloway, 1913 Late Ordovician Ashgill USA (Indiana)
  49. Heterotrypa obscura
  50. (Ulrich, 1883) Late Ordovician Caradoc USA (Ohio)
  51. Heterotrypa ovata
  52. Astrova, 1957 Late Silurian Ludlovian Russia
  53. Heterotrypa pajkhoyensis
  54. (Astrova, 1948) Middle Ordovician Russia
  55. Heterotrypa parvulipora
  56. Ulrich & Bassler, 1904 Late Ordovician Caradoc USA (Tennessee, Kentucky)
  57. Heterotrypa patera
  58. Coryell, 1921 Late Ordovician Caradoc USA (Tennessee, Alabama)
  59. Heterotrypa pauca
  60. Perry, 1962 Late Ordovician Caradoc USA (Iowa, Illinois)
  61. Heterotrypa paupera
  62. (Ulrich, 1883) Late Ordovician Caradoc USA (Ohio)
  63. Heterotrypa peripherilata
  64. Fan & Hu, 1985 Late Silurian China (Yunnan)
  65. Heterotrypa? perplexa
  66. Perry & Hattin, 1960 Silurian, Wenlockian USA (Indiana)
  67. Heterotrypa pontensis
  68. Ross, 1961 Early Devonian Australia (Vic.)
  69. Heterotrypa praenuntia
  70. (Ulrich, 1893) Late Ordovician Caradoc USA Minnesota, Estonia, Canada
  71. Heterotrypa praenuntia alexandria
  72. (Fritz, 1957) Ordovician Caradoc=Sandbian-Katian Canada Ontario
  73. Heterotrypa praenuntia echinata
  74. (Ulrich, 1893) Late Ordovician Caradoc USA, Canada
  75. Heterotrypa praenuntia multipora
  76. (Ulrich, 1893) Late Ordovician Caradoc USA (Minnesota)
  77. Heterotrypa praenuntia naevigera
  78. (Ulrich, 1893) Late Ordovician Caradoc USA, Estonia
  79. Heterotrypa praenuntia simplex
  80. (Ulrich, 1893) Late Ordovician Caradoc USA, Estonia
  81. Heterotrypa pretiosa
  82. Modzalevskaya, 1968 Silurian Ludlovian Russia
  83. Heterotrypa qujingensis
  84. Fan & Hu, 1985 Late Silurian China (Yunnan)
  85. Heterotrypa? ramosa
  86. (Owen, 1960) Late Silurian Ludlovian England
  87. Heterotrypa rapinae
  88. Pedder, 1971 Early Devonian Pragian Australia (NSW)
  89. Heterotrypa ridleyana
  90. (Coryell, 1921) Late Ordovician Caradoc USA (Tennessee, Alabama)
  91. Heterotrypa ringerikensis
  92. Ernst & Nakrem, 2015 Silurian Wenlock Norway
  93. Heterotrypa robusta
    (Homonym?)
  94. (Foord, 1884) Ordovician Ashgill=Katian-Hirnantian Canada Ontario
  95. Heterotrypa robusta
  96. Dyer, 1925 Late Ordovician Ashgill
    Silurian
    Canada
    Mongolia
  97. Heterotrypa rugosa
  98. Marintsch, 1998 Middle Ordovician USA (Tennessee)
  99. Heterotrypa semisphaerica
  100. Pushkin, 1976 Ordovician Llandeilo Belarus
  101. Heterotrypa simplex
  102. Dyer, 1925 Late Ordovician Ashgill Canada
  103. Heterotrypa simplex maculosa
  104. Dyer, 1925 Late Ordovician Ashgill Canada
  105. Heterotrypa singularis
  106. Ulrich, 1890 Late Ordovician Ashgill USA (Illinois, Wisconsin, Indiana)
  107. Heterotrypa sladei
  108. Buttler, 1991 Late Ordovician Ashgill Wales
  109. Heterotrypa solitaria
  110. Ulrich, 1883 Late Ordovician Caradoc USA, Canada
  111. Heterotrypa solitaria
    (Homonym)
  112. (Dyer, 1925) Late Ordovician Ashgill Canada
  113. Heterotrypa spinosa
  114. Kopajevich, 1984 Silurian Landovery-Wenlockian Mongolia
  115. Heterotrypa subfrondosa
  116. (Cumings, 1902) Late Ordovician Ashgill USA (Indiana)
  117. Heterotrypa subramosa
  118. (Ulrich, 1879) Late Ordovician Ashgill USA, Canada
  119. Heterotrypa subtrentonensis
  120. Marintsch, 1998 Middle Ordovician USA (Tennessee)
  121. Heterotrypa taffi
  122. Loeblich, 1942 Late Ordovician Caradoc USA (Oklahoma)
  123. Heterotrypa tenuis
  124. Astrova, 1959 Silurian Ludlovian Russia
  125. Heterotrypa trentonensis
  126. (Ulrich, 1883) Late Ordovician Caradoc USA, Canada
  127. Heterotrypa ulrichi
  128. (Nicholson, 1881) Late Ordovician Caradoc USA
  129. Heterotrypa undata
    undulata?
  130. Stewart, 1922 Devonian ?Pragian USA
  131. Heterotrypa wilmingtonensis
  132. (Ulrich & Bassler, 1904) Late Ordovician USA, Russia, Mongolia
  133. Heterotrypa? zubovkaensis
  134. Modzalevskaya, 1968 Silurian Wenlock Russia

This information is based upon various sources, but no personal knowledge. I am indebted to the late Alan Horowitz for his compilation, and to Dr. J. Pachut.
I would be grateful for corrections to, or critical reviews of this information. This is not a 'final' version (an impossible ideal), but a working document.

Heterotrypa praenuntia has also been referred to Dekayia, which presumably would reassign all the subspecies to the latter genus.

Dekayella (=Heterotrypa) at University of Georgia.
Dekayella (=Heterotrypa) from Nashville region: at University of Georgia.
Heterotrypa at University of Georgia.
Heterotrypa from Nashville region: at University of Georgia.

Heterotrypa subfrondosa reconstructions at DryDredgers.org.


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Edited by Phil Bock
Modified on 3/07/2015
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