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International Bryozoology Association
PRESIDENT Peter Hayward
J.R.P. Ross
COUNCIL MEMBERS 1995-2001 P. Bock
J. Cancino
J.-L. d'Hondt
M. Key Jr.
J. Scholtz
T.S. Wood
COUNCIL MEMBERS 1998-2004 A. Herrera
P. Moissette
A. Ostrovsky
J.A. Todd
J.E. Winston
E. Wöss


Micha Bayer has finally finished his Ph.D. at St Andrews. He has now started his new job at the Botanic Garden in Edinburgh. His project is on the development of an automatic computer-based identification system for diatoms. This is a large collaborative project in conjunction with seven European universities.

Mark Ward is presently working at Nettlecombe Court (back with the Field Studies Council). It is largely a teaching position but he is hoping to have more chances to get out into the field and look at bryozoans along the neglected Exmoor coast.


New Members

Dr. Sergey V. Dobretsov has been studying marine biofouling at die Biological Institute of the St Petersburg State University for more than 10 years. The main aims of the investigations at the White Sea are to study the formation and development of marine biofouling communities and the problem of non-toxic ecologically sensitive defense from biofouling. Recently research on the mechanisms of the settlement and substrate choice in different invertebrate larvae (bryozoans, hydroids, bivalves and barnacles) is in progress.

Natalia Shunatova is studying invertebrate zoology at St Petersburg State University. She is working on individual and colonial behaviour of bryozoans from the White Sea and at the moment her Master's Thesis on the anatomy and ultrastructure of tentacles of marine bryozoans is in preparation.

Irina Antipenko is studying anatomy and development in Phylactolaemata at St Petersburg State University. She is primarily interested in morphology, functioning of the neuro-muscular system and behaviour of Cristatella mucedo. She has also worked on the different aspects of budding and statoblastogenesis in Plumatella repens.

Jo Snell
is Paul Taylor's new Ph.D. student. She is starting work on the "Palaeobiology of Silurian Bryozoans", focusing mainly on the Wenlock bryozoans of the West Midlands and the Welsh Borderlands. Jo will be based at the University of Birmingham most of the time, supervised by Dr M.P. Smith.



Alan S. Horowitz [1930-1999]

Dr Alan S. Horowitz, 68, died on February 18, 1999 in Bloomington, Indiana, USA. He was curator of Paleontology, an emeritus Senior Research Scientist, and part-time Professor of Geology at Indiana University. Alan was born on June 12, 1930 in Ashland, Kentucky. He graduated Phi Beta Kappa from Washington and Lee University in 1952 and completed a Masters degree on igneous rocks of Greenland at The Ohio State University in 1953 and a Doctoral degree on the fauna of the Late Mississippian Glen Dean Formation at Indiana University in 1957.

Alan was a research geologist with Marathon Oil Company in Denver, Colorado, from 1957 to 1965, where he served as paleontologist with a team studying carbonate rocks. He returned to Indiana University as Curator of Paleontology in 1966 and served in that capacity until his retirement in 1996.

An expert on the geology of Indiana and the south-central United States, Alan specialized in Late Mississippian (Chesterian) rocks and fossils He was an internationally known authority on fossil bryozoans and blastoids. He was author or co-author of 53 papers, 57 abstracts, 1 fieldtrip guidebook, 5 bulletins and monographs, and I book. He was a member of many scientific and scholarly organizations and was a recognized bibliophile, with a library exceeding 3000 books.

Alan was an active member of the International Bryozoology Association and regularly attended its conferences. He commonly collaborated with fellow geologists and paleontologists, including especially Joseph Pachut and James Stratton in his studies of bryozoans.

We have lost a major contributor to the study of bryozoan systematics, diversity, and paleobiology. However, Alan will be remembered even more for his friendship and genuine warmth toward his colleagues.

Joseph F. Pachut


11th IBA Conference, Panama, 1998

Abstract titles

Bader B. Growth rate and life span of different Cellaria sinuosa (Hassall) colonies from the temperate shelf off North Brittany (France).

Barnes D.K.A. Life patterns of the encrusting Antarctic Cheilostomatida.

Barnes D.K. A. Life patterns of erect Antarctic Cheilostomatida.

Barnes D.K.A. & Sanderson W.G. Latitudinal patterns in the colonisation of marine debris.

Bock P.E. & Cook P.L. Early astogeny of Adeona colonies.

Brumbaugh D.R. Molecular phylogenetics of Schizoporellidae: a preliminary report.

Orellana M.C. & Manriquez P. Physiological ecology of marine bryozoans in temperate shallow water.

Cheetham A.H. & Jackson J.B.C. Neogene history of cheilostome Bryozoa in Tropical America.

Coates A.G. Neogene geologic history of the Isthmus of Panama.

Cocito S., Ferdeghini F., Morri C. & Bianchi C.N. Living bryozoan buildups: Schizoporella errata (Waters 1848) (Cheilostoma, Ascophora) in N. W. Mediterranean.

Cook P.L. & Bock P.E. Occurrence of two colony growth phases in Dimetopia hirta Macgillivray from Australia.

Craig S.F. & Wasson I.C. Self/non-self recognition in the bryozoan Hippodiplosia insculpta.

d'Hondt J.-L. Discovery of some types of cheilostomatous Bryozoa described in 1816 and 1824 by J.-F.-F. Lamouroux.

Dick M.H., Williams L.P. & Coggeshall-Burr M. Use of 16s mitochondrial ribosomal DNA sequences to investigate sister-group relationships among gymnolaemate bryozoans.

El Safori Y. A. Paleoenvironmental implications of the Middle Eocene bryozoans in Egypt.

El Safori Y. A. Early Eocene bryozoans of the Farafra Oasis, Egypt.

Fortunato H., Budd A.F., Foster C.T., Jr. & Golden J. Bryozoans on the World-Wide Web: the Neogene marine biota of Tropical America (NMITA database).

Gordon D.P. Phylogeny of the Cheilostomatida.

Hageman S.H. & Lidgard S. Bryozoa data exchange.

Hakansson E. Free-living bryozoans from the Paleocene of Western Australia.

Harmelin J.G. Ecology of cave and cavity dwelling bryozoans.

Hillmer G. & Voigt E. New genera of cyclostome bryozoans from the Upper Cretaceous (Germany) and paleocological implications of their occurrence.

Horowitz A.S. & Pachut J.F. Bryozoan species diversity from the fossil record.

Jackson J.B.C. & Herrera C. A. Adaptation and constraint as determinants of zooid and ovicell size among encrusting cheilostome Bryozoa from opposite sides of the Isthmus of Panama.

Jones K.E., Marsh T.G. & Wood T.S. Surveying for phylactolaemate bryozoans by sieving lentic sites for their statoblasts.

Key, Jr. M.M., Jeffries W.B., Voris H.K. & Yang C.M. Bryozoan fouling pattern on the horseshoe crab Tachypleus gigas (Muller) from Singapore.

Larsen N. & Hakansson E. Latest Maastrichtian bryozoan mounds from Stevns Klint, Denmark.

Lidgard S. & Hageman S. Recognizing bryozoan species.

McCartney M.A. Sexual selection in bryozoans.

McKinney F.K. Colony sizes and occurrence patterns among Bryozoa encrusting individual pelecypod valves.

McKinney F.K. Phylloporinids and the phylogeny of the Fenestrida.

Moissette P. The use of Neogene bryozoans for a better understanding of the ecology of some recent speces.

Moyano G., H.I. Bryozoa from the Magellanic Continental Slope near Cap Horn: an unexpected collection.

Nebelsick J.H. Biological and physical controls of rounded celleporid bryozoan growth.

Nielsen C. The phylogenetic position of entoprocts and ectoprocts based on morphological and molecular data.

Occhipinti Ambrogi A. Biodiversity and environmental sum through die recent developments of the history of bryozoans in the Lagoon of Venice.

O'Dea A. & Okamura B. Intracolonial variation in cheilostome bryozoans as an indicator of seasonality in the Pliocene.

Okamura B. The metapopulation biology of freshwater bryozoans.

Porter J.S., Ryland J.S. & Carvalho G.R. Species identification in the genus Alcyonidium Lamoureux (Bryozoa: Ctenostomatida) using a molecular approach.

Porter J.S., Bloor P., Cadman P.S., Stokell B.L. & Ryland J.S. Intra- and interspecific variation in the tentacle number of species in the genus AIcyonidium (Ctenostomatida)de Patron S., Ryland J.S. & Porter J.S. Ecology of Fucus-dwelling Alcyonidium spp. (Ctenostomatida) and the effect of an oil spillage on a South Wales population.

Ross J.R.P., Hamedi M.A. & Wright A.J. Late Ordovician (Caradoc) bryozoans from the Kerman District, East-Central ham

Ryland J.S. Gonozooid placement and branching patterns in some species of Crisia (Cyclostomatida).

Sanderson W.G. & Thorpe J.P. Aspects of structure and function in the feeding of the Antarctic cheilostomate bryozoan Himantozoum antarcticum.

Santagata S. & Zimmer R. Comparative cell activity patterns from bryozoan coronate larvae.

Schafer P. Bryozoan community dynamics and the Holocene evolution of bryozoan carbonates in the northern Atlantic.

Schellenberger J.S.& Ross J.R.P. Antibacterial substances in two marine cheilostomates (Bryozoa), northwest Washington.

Scholz J. Fungal infection and bryozoan morphology.

Scholz J. & Rainier G. Bryostromatolites.

Spjeldnaes N. Cryptic bryozonas from West Africa.

Taylor P.D. Cyclostome systematics: characters, phylogeny and classification.

Todd J.A. The central role of ctenostomes in bryozoan phylogeny.

Vavra N. Some biogeographical aspects of bryozoan fauna from the Austrian Neogene.

Wasson K. An experimental approach to colonial sexual strategies: testing predictions of sex allocation theory.

Weedon M.J. & Taylor P.D. Skeletal ultrastructure of primitive cheilostome bryozoans.

Winston J.E. & Hayward P.J. Marine bryozoans of the northeast coast of the United States.

Wood T.S. & Wood L.J. Statoblast morphology in historic specimens of phylactolaemate bryozoans.

Wöss E.R. Colonization and development of freshwater bryozoan communities on artificial substrates in the Laxenburg Pond (Lower Austria).


Restricted Access at NHM

There will be restricted access to the Recent Bryozoa collections at the NHM from 1st January - 31st December 2001. This is due to the construction of a new budding to house the fluid preserved zoological collections. Further information will be passed on, as and when details emerge.



1st Larwood Symposium, Bristol

On Friday I 11th December 1998, the School of Biological Sciences, University of Bristol, in a nutshell: Aaron O'Dea hosted the first Larwood Symposium. Named in honour of the late, and greatly missed, Gilbert Larwood, this meeting was a chance for (mainly) British bryozoan workers to gather together and more or less informally exchange ideas about work in progress. With the main IBA meetings being spaced at about three years, (and so being potentially missable during a research studentship), and with funding for travel to far-flung places hard to come by, the meeting provided an opportunity for "local" bryozoan workers to meet each other and keep in touch with current research. The format was simple: four sessions of roughly 15-20 minute presentations plus discussion, interspersed with refreshment breaks and lunch. The second day was devoted to field-collecting in l ocal fossil sites. In effect it was like a mini-IBA meeting, without the pain of processing manuscripts or the conference fee!

The delegates were largely Britain-based, although there was also a very impressive Scandinavian contingent in the form of Eckart Hakansson, Nicholas Larsen and Nils Spjeldnaes. A more accurate account of proceedings (i.e. abstracts and timetable) can be found on the website at http:/www.bio.bris.ac.uk/research/markwilk/larwood.htm

After the much-needed coffee and brief welcoming speech by Aaron, Kevin Tilbrook kick-started the symposium by introducing us to difficulties of unraveling the tangled web of taxonomy of south-west Pacific bryozoans. Kevin and Peter Hayward are making great strides towards preparing a synopsis-style guide for these bryozoans. John Ryland and Joanne Porter followed with a discussion of the complexities of taxonomy of at least seven species of British nearshore Alcyonidium.

These ctenostomes live, like trolls, under bridges, and can be distinguished by morphological characters as well as genetic (RAPD) evidence. John Bishop told us about his reproduction experiments with the ascidian Diplosoma and Celleporella: work he's been doing with Pato Manriquez; and Roger Hughes. The wizardry of Pato in constructing experimental apparatus, introduced here, was one of the themes of the symposium. Both the ascidian and the bryozoan, John concluded, require conspecific allosperm to be present before egg growth can initiate.

Sandwiched between mid-morning refreshments and lunch were two talks on the freshwater phylactolaemate Cristatella mucedo. The first by Tristan Hatton-Ellis concentrated on the population genetic structure of the species in lakes in the Thames basin of southern England. Comparison of clones both within and among different lakes showed that C. mucedo has a dynamic metapopulation structure, with all sites providing genetic diversity. Joanna Freeland (with Beth Okamura and Les Noble) used microsatellite genetic data to show that a metapopulation of C. mucedo occurs in the UK and western/north-western Europe. The distribution apparently fits with the transport of statoblasts by migrating wildfowl.

Lunch for delegates provided a stern test for the kitchen of the local Pizza Express. This provided a good opportunity to chew the bryozoan fat together, and for me at least, to chat in Britain to some folk I'd only ever seen in New Zealand, Panama or Japan; and to meet others I'd never met anywhere.

Bryozoans have the curious habit of drawing out great passion and enthusiasm in certain individuals, and no one is more enthusiastic than Paul Whittlesea, who will travel the earth as an amateur to attend IBA meetings, He only had to cross England on this occasion to start off an afternoon session featuring fossils and hard-parts, with a discussion of the possible defensive role of marginal budding of sub-colonies of the cyclostome Plagioecia from the Late Campanian. Nils Spjeldnaes entered well into the spirit of the symposium by bringing a neat selection of Plio-Pleistocene bryozoan specimens from Rhodes for our perusal. He described some of this fascinating material from his numerous field excursions to this pleasant Mediterranean island. In the second joint presentation of the day, Mike Weedon introduced the many calcitic ultrastructural fabrics occurring in cyclostome bryozoan skeletons, before Paul Taylor showed how these fabrics have been used with other hard part characters to produce the first substantial phylogeny for cyclostomes. Needless to say many of the audience became very pale when Paul flashed up the data matrix of ones and noughts and threatened to go through each character . Our final Scandinavian of the day was Eckart Hakansson who described the enormous wealth of cheilostome species (over one thousand) in the dramatic Maastrichtian radiation of Europe. He talked us through some of the adaptive strategies that evolved at that time.

Coffee and tea was a prelude to one last big session. A twist on the traditional IBA talk was given by Anthony Comer, who introduced the process of "data mining" . This is apparently a new way of extracting valuable seams of information from complex data warehouses. The techniques will be developed to be applied to bryozoans and other aspects of marine biology. Ewan Hunter [bravely coped without his slides??] then talked about his research in Japan on the larvae of that most important of bryozoans Bugula neritina. In addition to its apparent magical healing properties, this cheilostome is one of the major marine foulers. Ewan, and his colleagues Katsehiko Shimizu, Keiju Okano and Nobuhiro Fusetani, have found that the best thing to stop larvae settling is the hormone serotonin; they are hoping this can have commercial applications. Roger Hughes, (with the help of Pato Manriquez: interpreting some extraordinary slides) described a number of experiments involving large quantities of sperm and Celleporella hyalina. The final talk of the day was left to Beth Okamura (and J. E. Eckman), who showed how 2-dimensional modelling of suspension feeding in bryozoans can be used to imply that resource allocation may have played a key role in the evolution of colony form of encrusting bryozoans.

The conference was wound up by a brief word from the IBA secretary Mary Spencer Jones, followed immediately by the consumption of generous quantities of wine. The first Larwood Symposium was, in my opinion, a great success. It was a very good opportunity to catch up with bryozoan colleagues in Britain and Scandinavia and their dynamic research. A substantial part of this success must be owing to the excellent organisational skills of Aaron O'Dea, who should be greatly praised.
[Lord knows what happened on the field trip?]

Mike Weedon


New Bibliographic list on WWW

Prof. Giampietro Braga would like to announce that a bibliographic list on Earth Science of Tre Venezie: Veneto-Trentino-Alto Adige (South Tyrol) and Friuli (about 20,000 items) will be available on the Internet. This is a joint project between the Department of Geology, Paleontology and Geophysics, Padova University and the Civic Museum of Rovereto. You will find more than 3000 items on the Bryozoa, all with key words and part of an abstract. For more detailed information, open the website http://www.museocivico.rovereto.tn.it or write to the museum curator at x_francofinotti@museocivico.rovereto.tn.it

G. Braga Home Page:


IBA Business Meeting

30th January 1997, Tupper Center at the Smithsonian Tropical Research Institute, Balboa, Panama

1. At 3:50 p.m. Peter Hayward called the meeting to order.

2. Peter Hayward thanked Jeremy Jackson, Amalia Herrera, and all the speakers for making the 11th IBA such a productive conference. He then presented Amalia with an autographed conference volume from the 10th IBA Conference in New Zealand

3. Patrick Wyse Jackson gave a presentation on the 12th IBA conference to be held 16-21 July 2001 at the Department of Geology at Trinity College in Dublin, Ireland. Patrick will be the host. The conference will consist of four days of bryozoan talks, a one day mid-conference field trip, and a one day symposium on Saturday on the history of bryozoology. There win be a six day zoological preconference field trip 9-14 July and a six day geological post-conference field trip 23-28 July. Accommodations will be at Trinity College and will be about 30 Eire pounds/day. Registration will be about $450. The volume will probably be published by the Department of Geology at Trinity College.

4. Juan Cancino invited the IBA to hold its 13th IBA Conference in the 2nd or 3rd week of January 2004 in Concepcion, Chile. The organising committee includes Juan Cancino, Hugo Moyano, and Maria Cristina Orellana. The proposal was moved, seconded, and approved.

5. Peter Hayward observed the death of Gilbert Larwood who died in 1997. Paul Taylor gave a brief summary of Gilbert's career and contributions to the IBA. Peter announced that the 11th IBA volume would be dedicated to Gilbert. He proposed the establishment of the "IBA New Investigator Presentation Award". The proposal was moved, seconded, and approved.

6. Peter Hayward reported the successful housing of Professor Voigt's bryozoan collection in the Senckenberg Museum in Frankfurt. He proposed that the IBA send a letter of support for employing a curatorial assistant to oversee the collection. The proposal was moved, seconded, and approved.

7. Peter Hayward observed the death of Thomas Bolton in 1997. He gave a brief summary of Tom's career.

8. Peter Hayward read Mary Spencer Jones' treasurer's report. Acceptance of this report was moved, seconded, and approved. Peter announced that dues for 1998-2001 will be kept at $30/3 yr for regular members and $15/3 yr for student members. In 1997 we had 250 members.

9. Peter Hayward nominated Mary Spencer Jones for another three year term as Secretary/Treasurer. This was moved, seconded, and approved.

10. Phil Bock gave a presentation on the Bryozoa web page. He proposed that we publish the annual newsletter in hardcopy and on the web page. This proposal was moved seconded, and approved.

11. Peter Hayward reported that the six year terms of the following council members had ended: Bigey, Hageman, Hayward, Vavra, Wyse Jackson, and Ziko. The Executive Council nominated the following new council members: Herrera, Moissette, Ostrovsky, Todd, Winston, Woess. This new slate of council members was moved, seconded, mid approved. The following are the continuing council members: Bock, Cancino, d'Hondt, Key, Scholtz, Wood.

12. June Ross announced the Presidential Nominating Committee's recommendation for the next president be Ken McKinney. This was moved, seconded, and approved.

13. There being no further business, Peter Hayward closed the meeting at 4:32 p.m.

Respectfully submitted,
Marcus Key

[MSJ - As I was unable to attend the meeting in Panama, due to field work. Marcus kindly took the minutes.]