Smittina Norman, 1903

Type species: Lepralia landsborovii Johnston, 1847

Synonyms Author Type species
Grayporella Annoscia, 1969  
Levinseniula Cossman, 1920  
Marsillea Neviani, 1895 Millepora cervicornis Pallas, 1766
Porelloides Hayward, 1979 Cellepora laevis Fleming, 1828
Smittia (Homonym) Hincks, 1879 Lepralia landsborovii Johnston, 1847

This genus includes the species:

  1. Smittina abditavicularis
  2. Rogick, 1956 Recent Antarctica WoRMS 174362
  3. Smittina abyssicola
  4. (Harmer, 1957) Recent Indonesia WoRMS 470519
  5. Smittina acicularis
  6. Figuerola, Gordon & Cristobo, 2018 Recent Southwest Atlantic WoRMS 1055891
  7. Smittina acuminata
  8. (Ridley, 1881) Recent S. Atlantic WoRMS 470553
  9. Smittina affinis
  10. (Hincks, 1862) Recent NE Atlantic WoRMS 111549
  11. Smittina alata
  12. d'Hondt, 1986 Recent New Caledonia WoRMS 470500
  13. Smittina alticollarita
  14. Rogick, 1956 Recent Antarctica WoRMS 174364
  15. Smittina altirostris
  16. Osburn, 1952 Recent Pacific Alaska WoRMS 470539
  17. Smittina anecdota
  18. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 174366
  19. Smittina antarctica
  20. (Waters, 1904) Recent Antarctica WoRMS 174368
  21. Smittina arctica
  22. (Norman, 1894) Recent North Atlantic, Arctic WoRMS 853628
  23. Smittina areolata
  24. (O'Donoghue & O'Donoghue, 1923) Recent NE Pacific Canada WoRMS 853909
  25. Smittina asymmetrica
  26. Gordon & d'Hondt, 1997 Recent S Pacific New Caledonia WoRMS 470518
  27. Smittina? aviculifera
  28. (Canu, 1912) Paleogene Eocene Lutetian France  
  29. Smittina azorensis
  30. (Jullien, 1903) Recent Atlantic Azores WoRMS 470549
  31. Smittina baccata
  32. Canu & Bassler, 1930 Recent Mediterranean Tunisia WoRMS 470544
  33. Smittina bassleri
  34. Cipolla, 1928 Neogene Miocene Libya  
  35. Smittina bella
  36. (Busk, 1860) Recent NE Atlantic WoRMS 111550
  37. Smittina beringia
  38. Kluge, 1952 Recent Arctic WoRMS 470535
  39. Smittina? bidentata
  40. Androsova, 1958 Recent NW Pacific Sakhalin WoRMS 853804
  41. Smittina? brevis
  42. Canu & Bassler, 1919 Neogene Pliocene Jamaica  
  43. Smittina canavarii
  44. (Neviani, 1900) Neogene Miocene-Pliocene Italy, Spain, Algeria, Morocco  
  45. Smittina cervicornis
  46. (Pallas, 1766) Recent
    Paleogene-Neogene
    N Atlantic, Mediterranean
    Europe
    WoRMS 111551
  47. Smittina chilensis
  48. Moyano, 1991 Recent Chile WoRMS 470515
  49. Smittina colletti
  50. (Jullien, 1903) Recent NE Atlantic WoRMS 111553
  51. Smittina confluens
  52. (Reuss, 1864) Paleogene Oligocene Chattian Germany  
  53. Smittina confusa
  54. d'Hondt, 1978 Recent Mediterranean Tunisia WoRMS 1043829
  55. Smittina cophia
  56. Canu & Bassler, 1920 Paleogene Oligocene, Rupelian USA Alabama  
  57. Smittina cordata
  58. Osburn, 1952 Recent USA California, Gulf of Mexico WoRMS 470541
  59. Smittina coronata
  60. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA N. Carolina  
  61. Smittina crepidula
  62. Brown, 1952 Paleogene Oligocene Rupelian New Zealand  
  63. Smittina cribraria
  64. (MacGillivray, 1886) Recent
    Neogene Miocene Langhian
    Australia WoRMS 470556
  65. Smittina crystallina
  66. (Norman, 1867) Recent N Atlantic WoRMS 111554
  67. Smittina ctenocondyla
  68. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 470503
  69. Smittina curta
  70. Canu, 1926 Paleogene Eocene Priabonian France  
  71. Smittina curvirostrata
  72. De Blauwe & Gordon, 2014 Recent Antarctic - Weddell Sea WoRMS 470503
  73. Smittina cylindrica
  74. (Canu, 1908) Paleogene Eocene Lutetian France  
  75. Smittina deangelisi
  76. (Neviani, 1905) Paleogene Eocene Spain  
  77. Smittina? decipiens
  78. (Jullien, 1903) Recent Northeast Atlantic France WoRMS 853910
  79. Smittina depressa
  80. (MacGillivray, 1895) Neogene Miocene Langhian SE Australia  
  81. Smittina derwiesi
  82. (Canu, 1908) Paleogene Eocene Lutetian France, England, Belgium  
  83. Smittina dieuzeidei
  84. Gautier, 1955 Recent N Atlantic WoRMS 470536
  85. Smittina diffidentia
  86. Hayward & Thorpe, 1989 Recent Antarctica WoRMS 174441
  87. Smittina diplostoma
  88. (Philippi, 1843) Paleogene Oligocene Germany  
  89. Smittina directa
  90. (Waters, 1904) Recent Antarctica WoRMS 174372
  91. Smittina discoidea
  92. Canu & Bassler, 1923 Neogene Pleistocene USA California  
  93. Smittina eagari
  94. Brown, 1958 Neogene Miocene SE Australia  
  95. Smittina ectoproctolitica
  96. Moyano, 1982 Recent SW Atlantic WoRMS 470498
  97. Smittina emiliana
  98. Pizzaferri, 2004 Neogene Pliocene-Pleistocene
    Zanclean, Piacenzian
    Italy  
  99. Smittina euparypha
  100. Marcus, 1921 Recent SE Pacific WoRMS 470548
  101. Smittina excentrica
  102. (Canu, 1912) Paleogene Eocene Lutetian? France  
  103. Smittina excertiaviculata
  104. Rogick, 1956 Recent Antarctica WoRMS 470486
  105. Smittina? exclusa
  106. Harmer, 1957 Recent Indonesia WoRMS 470530
  107. Smittina exigua
  108. Canu & Bassler, 1920 Paleogene Eocene Priabonian SE USA  
  109. Smittina fallax
  110. (Jullien, 1903) Recent NE Atlantic WoRMS 470555
  111. Smittina ferruginea
  112. Hayward & Cook, 1983 Recent S Africa WoRMS 470496
  113. Smittina fontensis
  114. Julien, 1940 Neogene Miocene Langhian France  
  115. Smittina forata
  116. Canu & Bassler, 1929 Paleogene Eocene Lutetian Belgium  
  117. Smittina forticula
  118. Winston, 2016 Recent Northwest Atlantic USA Florida WoRMS 868521
  119. Smittina fragaria
  120. Moyano, 1983 Recent Chile WoRMS 470520
  121. Smittina glebula
  122. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 174443
  123. Smittina grandicella
  124. Canu & Bassler, 1923 Neogene Pleistocene USA California  
  125. Smittina grandifossa
  126. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA North Carolina  
  127. Smittina? granulata
  128. (Canu, 1908) Neogene Miocene Argentina  
  129. Smittina granulosa
  130. Canu & Bassler, 1920 Paleogene Oligocene Rupelian USA Alabama  
  131. Smittina hanaishiensis
  132. Hayami, 1975 Neogene Pliocene Japan  
  133. Smittina hatsushima
  134. Okada & Mawatari, 1937 Recent Japan WoRMS 470543
  135. Smittina humilis
  136. Gontar, 2008 Recent Antarctic WoRMS 470559
  137. Smittina impellucida
  138. Gontar, 2008 Recent Antarctic WoRMS 470560
  139. Smittina imragueni
  140. Matsuyama, Titschack, Baum & Freiwald, 2015 Recent East Atlantic Mauritania WoRMS 853191
  141. Smittina inarmata
  142. Brown, 1958 Neogene Miocene SE Australia  
  143. Smittina incernicula
  144. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 174378
  145. Smittina insulata
  146. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 470504
  147. Smittina invisitata
  148. Sokurov, 1975 Paleogene Paleocene Kazakhstan  
  149. Smittina isabelae
  150. Souto, Berning & Ostrovsky, 2016 Recent Northeast Atlantic Spain WoRMS 868425
  151. Smittina jacobensis
  152. (Busk, 1884) Recent S Atlantic WoRMS 470505
  153. Smittina jacquelinae
  154. Moyano, 1983 Recent Chile WoRMS 470521
  155. Smittina jordii
  156. Reverter Gil & Fernandez Pulpeiro, 1999 Recent NE Atlantic WoRMS 111557
  157. Smittina jullieni
  158. Moyano, 1983 Recent SW Atlantic WoRMS 470506
  159. Smittina kobjakovae
  160. Androsova, 1958 Recent NW Pacific WoRMS 470531
  161. Smittina kukuiula
  162. Soule & Soule, 1973 Recent Hawai'i WoRMS 470497
  163. Smittina kussakini
  164. Gontar, 1993 Recent NW Pacific WoRMS 470516
  165. Smittina labiatula
  166. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA North Carolina  
  167. Smittina landsborovii
  168. (Johnston, 1847) Recent Europe WoRMS 111558
  169. Smittina lebruni
  170. (Waters, 1905) Recent Atlantic WoRMS 470494
  171. Smittina lecointrei
  172. Buge, 1961 Neogene Miocene Tortonian France  
  173. Smittina? leda
  174. (d'Orbigny, 1852) Paleogene Eocene Lutetian France  
  175. Smittina leptodentata
  176. Hayward & Thorpe, 1990 Recent S Atlantic WoRMS 470507
  177. Smittina lobata
  178. (Busk, 1859) Neogene Miocene-Pliocene England, France, Portugal  
  179. Smittina maccullochae
  180. Osburn, 1952 Recent
    Neogene Pleistocene
    NE Pacific
    USA California
    WoRMS 470540
  181. Smittina? macgillivrayi
  182. (Maplestone, 1902) Neogene Miocene Australia Victoria.  
  183. Smittina maleposita
  184. Canu & Bassler, 1923 Neogene Pleistocene USA Florida  
  185. Smittina malleolus
  186. (Hincks, 1884) Recent South Pacific - Australia Bass Strait WoRMS 111560
  187. Smittina malouinensis
  188. (Jullien, 1888) Recent S Atlantic WoRMS 470552
  189. Smittina marionensis
  190. (Busk, 1854) Recent S Indian O. WoRMS 470493
  191. Smittina messiniensis
  192. el Hajjaji, 1992 Neogene Miocene Messinian Morocco  
  193. Smittina microtheca
  194. Canu & Lecointre, 1930 Neogene Miocene Tortonian France  
  195. Smittina migottoi
  196. Vieira, Gordon, Souza & Haddad, 2010 Recent West Atlantic - Brazil WoRMS 468327
  197. Smittina minima
  198. Powell, 1967 Recent New Zealand WoRMS 470528
  199. Smittina minuscula
  200. (Smitt, 1868) Recent Arctic WoRMS 111561
  201. Smittina molarifera
  202. Moyano, 1982 Recent Chile WoRMS 470499
  203. Smittina monacha
  204. (Jullien, 1888) Recent S Atlantic WoRMS 470508
  205. Smittina mucronata
  206. (Smitt, 1868) Recent Arctic, Canada, Japan WoRMS 470508
  207. Smittina muliebris
  208. Kluge, 1962 Recent Arctic WoRMS 423756
  209. Smittina multanguloporata
  210. Gontar, 2008 Recent Antarctic WoRMS 470558
  211. Smittina nitidissima
  212. (Hincks, 1880) Recent Atlantic, world-wide? WoRMS 220640
  213. Smittina nodai
  214. Hayami, 1975 Neogene Pliocene Japan  
  215. Smittina normani
  216. Aristegui, 1988 Recent E Atlantic WoRMS 470502
  217. Smittina obicullata
  218. Rogick, 1956 Recent Antarctica WoRMS 174380
  219. Smittina obicullatoidea
  220. Liu & Hu, 1991 Recent Antarctica WoRMS 470526
  221. Smittina oblita
  222. Lopez Gappa, 2002 Recent Argentina WoRMS 387454
  223. Smittina obscura
  224. (MacGillivray, 1891) Recent Australia WoRMS 470550
  225. Smittina oculata
  226. (MacGillivray, 1883) Recent Australia WoRMS 470551
  227. Smittina orbavicularia
  228. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA Carolinas  
  229. Smittina ordinata
  230. MacGillivray, 1895 Neogene Miocene SE Australia  
  231. Smittina osburni
  232. Hayami, 1975 Neogene Miocene-Pliocene Japan  
  233. Smittina ovirotula
  234. Soule, Soule & Chaney, 1995 Recent NE Pacific California WoRMS 470514
  235. Smittina palisada
  236. Gordon, 1989 Recent New Zealand WoRMS 470492
  237. Smittina papillifera
  238. (MacGillivray, 1869) Recent SE Australia WoRMS 470485
  239. Smittina? parviovicellosa
  240. Canu & Bassler, 1935 Paleogene Oligocene- Neogene Miocene Australia Victoria.  
  241. Smittina personata
  242. (Hincks, 1884) Recent SE Australia WoRMS 470491
  243. Smittina pileata
  244. (Waters, 1904) Recent Antarctica WoRMS 174383
  245. Smittina pliofistulata
  246. Hayward & Thorpe, 1990 Recent SW Atlantic WoRMS 470509
  247. Smittina pocilla
  248. Hayward & Thorpe, 1990 Recent SW Atlantic WoRMS 174445
  249. Smittina porelloides
  250. (Canu & Bassler, 1920) Paleogene Eocene Priabonian USA, Hungary  
  251. Smittina porosa
  252. Canu & Bassler, 1928 Recent East Atlantic Morocco WoRMS 1043830
  253. Smittina porrigens
  254. (Reuss, 1865) Paleogene Oligocene Rupelian Germany  
  255. Smittina portiuscula
  256. Hayward & Thorpe, 1990 Recent SW Atlantic WoRMS 470510
  257. Smittina pouyetae
  258. el Hajjaji, 1992 Neogene Miocene Messinian Morocco  
  259. Smittina protrusa
  260. Powell, 1967 Recent, Pleistocene Australia WoRMS 470529
  261. Smittina pseudoacutirostris
  262. Gostilovskaya, 1957 Recent Arctic WoRMS 423758
  263. Smittina? pseudocompressa
  264. Androsova, 1958 Recent NW Pacific Sakhalin WoRMS 470534
  265. Smittina? pulchra
  266. Androsova, 1958 Recent NW Pacific WoRMS 470532
  267. Smittina? punctifera
  268. (Canu, 1908) Neogene Miocene Argentina  
  269. Smittina puncturata
  270. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA North Carolina  
  271. Smittina? pupa
  272. Canu & Bassler, 1920 Paleogene Oligocene Rupelian USA Alabama  
  273. Smittina purpurea
  274. (Hincks, 1881) Recent Australia WoRMS 470490
  275. Smittina remotorostrata
  276. (Canu & Bassler, 1928) Recent
    Neogene Miocene Messinian
    Mediterranean, NE Atlantic
    Morocco
    WoRMS 111562
  277. Smittina reptans
  278. (Waters, 1904) Recent Antarctic WoRMS 853629
  279. Smittina reticuloides
  280. Canu & Bassler, 1920 Paleogene Oligocene Rupelian USA Alabama  
  281. Smittina retifrons
  282. Osburn, 1952 Recent N Pacific Alaska WoRMS 470538
  283. Smittina rogickae
    (Homonym)
  284. Hayward & Taylor, 1984 Recent
    Neogene Pleistocene
    Antarctica WoRMS 174386
  285. Smittina rosacea
  286. Powell, 1967 Recent New Zealand WoRMS 470487
  287. Smittina rosefieldensis
  288. McGuirt, 1941 Paleogene Oligocene Rupelian USA Louisiana  
  289. Smittina salsa
  290. Pizzaferri, 2004 Neogene Pliocene-Pleistocene Italy  
  291. Smittina scrupea
  292. Hayward & Taylor, 1984 Neogene Pleistocene Antarctica WoRMS 470524
  293. Smittina seguenzai
  294. (Reuss, 1869) Paleogene Eocene Priabonian Italy  
  295. Smittina sextapuncta
  296. (Canu, 1912) Paleogene Eocene Lutetian France  
  297. Smittina sigillata
  298. (Jullien, 1888) Recent South Atlantic Falklands WoRMS 853911
  299. Smittina sitella
  300. Hayward & Cook, 1983 Recent S Africa WoRMS 470495
  301. Smittina smitti
  302. (Kirchenpauer, 1874) Recent Arctic WoRMS 111564
  303. Smittina smittiana
  304. (Busk, 1884) Recent S Atlantic WoRMS 470511
  305. Smittina smittiella
  306. Osburn, 1947 Recent West Atlantic - Caribbean WoRMS 422353
  307. Smittina sordida
  308. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA Georgia  
  309. Smittina spathulifera
  310. (Hincks, 1884) Recent NE Pacific Canada WoRMS 470554
  311. Smittina spiraminifera
  312. Gordon, 1984 Recent SW Pacific Kermadec WoRMS 470488
  313. Smittina stigmatophora
  314. (Busk, 1884) Recent SW Atlantic WoRMS 470512
  315. Smittina stricta
  316. Vigneaux, 1949 Neogene Miocene Tortonian? France  
  317. Smittina subcordata
  318. Gontar, 1993 Recent NW Pacific WoRMS 470517
  319. Smittina terraenovae
  320. Brown, 1954 Neogene Pliocene
    Recent
    New Zealand WoRMS 470537
  321. Smittina thompsoni
  322. Kluge, 1955 Recent Arctic WoRMS 465715
  323. Smittina torques
  324. Powell, 1967 Recent New Zealand WoRMS 470489
  325. Smittina triangularis
  326. Duvergier, 1921 Neogene Miocene Burdigalian France  
  327. Smittina tuberosa
  328. Kluge, 1952 Recent N Pacific WoRMS 470525
  329. Smittina tubulata
  330. (Gabb & Horn, 1862) Paleogene Oligocene Rupelian USA Alabama  
  331. Smittina umbilicata
  332. (Römer, 1863) Oligocene Miocene Europe  
  333. Smittina undulimargo
  334. Moyano, 1983 Recent Chile WoRMS 470522
  335. Smittina unicus
  336. Tilbrook, 2006 Recent Solomon Islands WoRMS 470557
  337. Smittina uruguayensis
  338. d'Hondt, 1981 Recent SW Atlantic WoRMS 470501
  339. Smittina? uschakowi
  340. Androsova, 1958 Recent NW Pacific WoRMS 470533
  341. Smittina uttleyi
  342. Brown, 1958 Neogene Miocene SE Australia  
  343. Smittina vaciva
  344. (Jullien, 1882) Recent Northeast Atlantic France Bay of Biscay old WoRMS 1043831
  345. Smittina variavicularis
  346. Mawatari, 1974 Recent Japan WoRMS 470527
  347. Smittina veleroa
  348. Soule, Soule & Chaney, 1995 Recent NE Pacific California WoRMS 470513
  349. Smittina volcanica
  350. Moyano, 1983 Recent Chile WoRMS 470523

The list below is of species identified as "Smittia" which appear to have no subsequent revision. They may belong to Smittina, but equally may be species of Parasmittina or Smittoidea.

  1. "Smittia" gemmata
  2. Jullien, 1903 Recent Northeast Atlantic France Gulf of Gascony
  3. "Smittia" grimaldii
  4. Jullien, 1903 Recent Northeast Atlantic France Gulf of Gascony
  5. "Smittia" immersa
  6. Jullien, 1903 Recent Northeast Atlantic France Gulf of Gascony
  7. "Smittia" incisa
  8. Canu, 1904 non (Reuss, 1873) Neogene Argentina
  9. "Smittia" levinseni
  10. Nordgaard, 1903 Recent Arctic
  11. "Smittia" miniacea
  12. Jullien, 1882 Recent Northeast Atlantic France Bay of Biscay
  13. "Smittia" reticulata nitida
  14. MacGillivray, 1895 Neogene Miocene Langhian Australia Victoria
  15. "Smittia" robusta
  16. Jullien, 1903 Recent Northwest Atlantic Grand Banks
  17. "Smittia" schlumbergeri
  18. Jullien, 1903 Recent Northwest Atlantic Grand Banks
  19. "Smittia" subtorquata
  20. Canu, 1904 Neogene Argentina
  21. "Smittia" zuccari
  22. Neviani, 1895 Neogene Pliocene Italy
Smittina kukuiula Smittina nitidissima Smittina papillifera Smittina sp.1
Smittina kukuiula
(Thanks to Dr D. Soule)
Smittina nitidissima Smittina papillifera Smittina sp.1

This information is based upon various sources, but little personal knowledge. I am indebted to the late Alan Horowitz for his compilation, and to Dr. J. Pachut.
I would be grateful for corrections to, or critical reviews of this information. This is not a 'final' version (an impossible ideal), but a working document.

Harmer notebook - entry for Smittia
Harmer notebook - entry for Smittina

Smittina page at NMITA

Smittina bella at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina beringia at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina landsborovii at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina minuscula at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina mucronata at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina pseudoacutirostris at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina smitti at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina thompsoni at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina tuberosa at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittoidea glaciata at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittoidea reticulata at Atlas of Arctic Bryozoa - Piotr Kuklinski.

Smittina hanaishiensis at Pleistocene Bryozoa of Kuromatsunai, Japan.


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Edited by Phil Bock
Modified on 31/3/2017
Modified on 11/11/2017 (Five species added, also "Smittia" spp.)
Modified on 24/1/2018 (Added S. aciculata)
This URL is http://bryozoa.net/cheilostomata/smittinidae/smittina.html