Smittina Norman, 1903

Type species: Lepralia landsborovii Johnston, 1847

Synonyms Author Type species
Grayporella Annoscia, 1969  
Levinseniula Cossman, 1920  
Marsillea Neviani, 1895 Millepora cervicornis Pallas, 1766
Porelloides Hayward, 1979 Cellepora laevis Fleming, 1828
Smittia (Homonym) Hincks, 1879 Lepralia landsborovii Johnston, 1847

This genus includes the species:

  1. Smittina abditavicularis
  2. Rogick, 1956 Recent Antarctica WoRMS 174362
  3. Smittina abyssicola
  4. (Harmer, 1957) Recent Indonesia WoRMS 470519
  5. Smittina acicularis
  6. Figuerola, Gordon & Cristobo, 2018 Recent Southwest Atlantic WoRMS 1055891
  7. Smittina acuminata
  8. (Ridley, 1881) Recent S. Atlantic WoRMS 470553
  9. Smittina affinis
  10. (Hincks, 1862) Recent NE Atlantic WoRMS 111549
  11. Smittina alata
  12. d'Hondt, 1986 Recent New Caledonia WoRMS 470500
  13. Smittina alticollarita
  14. Rogick, 1956 Recent Antarctica WoRMS 174364
  15. Smittina altirostris
  16. Osburn, 1952 Recent Pacific Alaska WoRMS 470539
  17. Smittina anecdota
  18. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 174366
  19. Smittina antarctica
  20. (Waters, 1904) Recent Antarctica WoRMS 174368
  21. Smittina arctica
  22. (Norman, 1894) Recent North Atlantic, Arctic WoRMS 853628
  23. Smittina areolata
  24. (O'Donoghue & O'Donoghue, 1923) Recent NE Pacific Canada WoRMS 853909
  25. Smittina asymmetrica
  26. Gordon & d'Hondt, 1997 Recent S Pacific New Caledonia WoRMS 470518
  27. Smittina? aviculifera
  28. (Canu, 1912) Paleogene Eocene Lutetian France  
  29. Smittina azorensis
  30. (Jullien, 1903) Recent Atlantic Azores WoRMS 470549
  31. Smittina baccata
  32. Canu & Bassler, 1930 Recent Mediterranean Tunisia WoRMS 470544
  33. Smittina bassleri
  34. Cipolla, 1928 Neogene Miocene Libya  
  35. Smittina bella
  36. (Busk, 1860) Recent NE Atlantic WoRMS 111550
  37. Smittina beringia
  38. Kluge, 1952 Recent Arctic WoRMS 470535
  39. Smittina? bidentata
  40. Androsova, 1958 Recent NW Pacific Sakhalin WoRMS 853804
  41. Smittina? brevis
  42. Canu & Bassler, 1919 Neogene Pliocene Jamaica  
  43. Smittina canavarii
  44. (Neviani, 1900) Neogene Miocene-Pliocene Italy, Spain, Algeria, Morocco  
  45. Smittina cervicornis
  46. (Pallas, 1766) Recent
    Paleogene-Neogene
    N Atlantic, Mediterranean
    Europe
    WoRMS 111551
  47. Smittina chilensis
  48. Moyano, 1991 Recent Chile WoRMS 470515
  49. Smittina colletti
  50. (Jullien, 1903) Recent NE Atlantic WoRMS 111553
  51. Smittina confluens
  52. (Reuss, 1864) Paleogene Oligocene Chattian Germany  
  53. Smittina confusa
  54. d'Hondt, 1978 Recent Mediterranean Tunisia WoRMS 1043829
  55. Smittina cophia
  56. Canu & Bassler, 1920 Paleogene Oligocene, Rupelian USA Alabama  
  57. Smittina cordata
  58. Osburn, 1952 Recent USA California, Gulf of Mexico WoRMS 470541
  59. Smittina coronata
  60. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA N. Carolina  
  61. Smittina crepidula
  62. Brown, 1952 Paleogene Oligocene Rupelian New Zealand  
  63. Smittina cribraria
  64. (MacGillivray, 1886) Recent
    Neogene Miocene Langhian
    Australia WoRMS 470556
  65. Smittina crystallina
  66. (Norman, 1867) Recent N Atlantic WoRMS 111554
  67. Smittina ctenocondyla
  68. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 470503
  69. Smittina curta
  70. Canu, 1926 Paleogene Eocene Priabonian France  
  71. Smittina curvirostrata
  72. De Blauwe & Gordon, 2014 Recent Antarctic - Weddell Sea WoRMS 470503
  73. Smittina cylindrica
  74. (Canu, 1908) Paleogene Eocene Lutetian France  
  75. Smittina deangelisi
  76. (Neviani, 1905) Paleogene Eocene Spain  
  77. Smittina depressa
  78. (MacGillivray, 1895) Neogene Miocene Langhian SE Australia  
  79. Smittina derwiesi
  80. (Canu, 1908) Paleogene Eocene Lutetian France, England, Belgium  
  81. Smittina dieuzeidei
  82. Gautier, 1955 Recent N Atlantic WoRMS 470536
  83. Smittina diffidentia
  84. Hayward & Thorpe, 1989 Recent Antarctica WoRMS 174441
  85. Smittina diplostoma
  86. (Philippi, 1843) Paleogene Oligocene Germany  
  87. Smittina directa
  88. (Waters, 1904) Recent Antarctica WoRMS 174372
  89. Smittina discoidea
  90. Canu & Bassler, 1923 Neogene Pleistocene USA California  
  91. Smittina eagari
  92. Brown, 1958 Neogene Miocene SE Australia  
  93. Smittina ectoproctolitica
  94. Moyano, 1982 Recent SW Atlantic WoRMS 470498
  95. Smittina emiliana
  96. Pizzaferri, 2004 Neogene Pliocene-Pleistocene
    Zanclean, Piacenzian
    Italy  
  97. Smittina euparypha
  98. Marcus, 1921 Recent SE Pacific WoRMS 470548
  99. Smittina excentrica
  100. (Canu, 1912) Paleogene Eocene Lutetian? France  
  101. Smittina excertiaviculata
  102. Rogick, 1956 Recent Antarctica WoRMS 470486
  103. Smittina? exclusa
  104. Harmer, 1957 Recent Indonesia WoRMS 470530
  105. Smittina exigua
  106. Canu & Bassler, 1920 Paleogene Eocene Priabonian SE USA  
  107. Smittina ferruginea
  108. Hayward & Cook, 1983 Recent S Africa WoRMS 470496
  109. Smittina fontensis
  110. Julien, 1940 Neogene Miocene Langhian France  
  111. Smittina forata
  112. Canu & Bassler, 1929 Paleogene Eocene Lutetian Belgium  
  113. Smittina forticula
  114. Winston, 2016 Recent Northwest Atlantic USA Florida WoRMS 868521
  115. Smittina fragaria
  116. Moyano, 1983 Recent Chile WoRMS 470520
  117. Smittina glebula
  118. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 174443
  119. Smittina grandicella
  120. Canu & Bassler, 1923 Neogene Pleistocene USA California  
  121. Smittina grandifossa
  122. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA North Carolina  
  123. Smittina? granulata
  124. (Canu, 1908) Neogene Miocene Argentina  
  125. Smittina granulosa
  126. Canu & Bassler, 1920 Paleogene Oligocene Rupelian USA Alabama  
  127. Smittina hanaishiensis
  128. Hayami, 1975 Neogene Pliocene Japan  
  129. Smittina hatsushima
  130. Okada & Mawatari, 1937 Recent Japan WoRMS 470543
  131. Smittina humilis
  132. Gontar, 2008 Recent Antarctic WoRMS 470559
  133. Smittina impellucida
  134. Gontar, 2008 Recent Antarctic WoRMS 470560
  135. Smittina imragueni
  136. Matsuyama, Titschack, Baum & Freiwald, 2015 Recent East Atlantic Mauritania WoRMS 853191
  137. Smittina inarmata
  138. Brown, 1958 Neogene Miocene SE Australia  
  139. Smittina incernicula
  140. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 174378
  141. Smittina insulata
  142. Hayward & Thorpe, 1990 Recent Antarctica WoRMS 470504
  143. Smittina invisitata
  144. Sokurov, 1975 Paleogene Paleocene Kazakhstan  
  145. Smittina isabelae
  146. Souto, Berning & Ostrovsky, 2016 Recent Northeast Atlantic Spain WoRMS 868425
  147. Smittina jacobensis
  148. (Busk, 1884) Recent S Atlantic WoRMS 470505
  149. Smittina jacquelinae
  150. Moyano, 1983 Recent Chile WoRMS 470521
  151. Smittina jordii
  152. Reverter Gil & Fernandez Pulpeiro, 1999 Recent NE Atlantic WoRMS 111557
  153. Smittina jullieni
  154. Moyano, 1983 Recent SW Atlantic WoRMS 470506
  155. Smittina kobjakovae
  156. Androsova, 1958 Recent NW Pacific WoRMS 470531
  157. Smittina kukuiula
  158. Soule & Soule, 1973 Recent Hawai'i WoRMS 470497
  159. Smittina kussakini
  160. Gontar, 1993 Recent NW Pacific WoRMS 470516
  161. Smittina labiatula
  162. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA North Carolina  
  163. Smittina landsborovii
  164. (Johnston, 1847) Recent Europe WoRMS 111558
  165. Smittina lebruni
  166. (Waters, 1905) Recent Atlantic WoRMS 470494
  167. Smittina lecointrei
  168. Buge, 1961 Neogene Miocene Tortonian France  
  169. Smittina? leda
  170. (d'Orbigny, 1852) Paleogene Eocene Lutetian France  
  171. Smittina leptodentata
  172. Hayward & Thorpe, 1990 Recent S Atlantic WoRMS 470507
  173. Smittina lobata
  174. (Busk, 1859) Neogene Miocene-Pliocene England, France, Portugal  
  175. Smittina maccullochae
  176. Osburn, 1952 Recent
    Neogene Pleistocene
    NE Pacific
    USA California
    WoRMS 470540
  177. Smittina? macgillivrayi
  178. (Maplestone, 1902) Neogene Miocene Australia Victoria.  
  179. Smittina malleolus
  180. (Hincks, 1884) Recent South Pacific - Australia Bass Strait WoRMS 111560
  181. Smittina malouinensis
  182. (Jullien, 1888) Recent S Atlantic WoRMS 470552
  183. Smittina marionensis
  184. (Busk, 1854) Recent S Indian O. WoRMS 470493
  185. Smittina messiniensis
  186. el Hajjaji, 1992 Neogene Miocene Messinian Morocco  
  187. Smittina microtheca
  188. Canu & Lecointre, 1930 Neogene Miocene Tortonian France  
  189. Smittina migottoi
  190. Vieira, Gordon, Souza & Haddad, 2010 Recent West Atlantic - Brazil WoRMS 468327
  191. Smittina minima
  192. Powell, 1967 Recent New Zealand WoRMS 470528
  193. Smittina minuscula
  194. (Smitt, 1868) Recent Arctic WoRMS 111561
  195. Smittina molarifera
  196. Moyano, 1982 Recent Chile WoRMS 470499
  197. Smittina monacha
  198. (Jullien, 1888) Recent S Atlantic WoRMS 470508
  199. Smittina mucronata
  200. (Smitt, 1868) Recent Arctic, Canada, Japan WoRMS 470508
  201. Smittina muliebris
  202. Kluge, 1962 Recent Arctic WoRMS 423756
  203. Smittina multanguloporata
  204. Gontar, 2008 Recent Antarctic WoRMS 470558
  205. Smittina nitidissima
  206. (Hincks, 1880) Recent Atlantic, world-wide? WoRMS 220640
  207. Smittina nodai
  208. Hayami, 1975 Neogene Pliocene Japan  
  209. Smittina normani
  210. Aristegui, 1988 Recent E Atlantic WoRMS 470502
  211. Smittina obicullata
  212. Rogick, 1956 Recent Antarctica WoRMS 174380
  213. Smittina obicullatoidea
  214. Liu & Hu, 1991 Recent Antarctica WoRMS 470526
  215. Smittina oblita
  216. Lopez Gappa, 2002 Recent Argentina WoRMS 387454
  217. Smittina obscura
  218. (MacGillivray, 1891) Recent Australia WoRMS 470550
  219. Smittina oculata
  220. (MacGillivray, 1883) Recent Australia WoRMS 470551
  221. Smittina orbavicularia
  222. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA Carolinas  
  223. Smittina ordinata
  224. MacGillivray, 1895 Neogene Miocene SE Australia  
  225. Smittina osburni
  226. Hayami, 1975 Neogene Miocene-Pliocene Japan  
  227. Smittina ovirotula
  228. Soule, Soule & Chaney, 1995 Recent NE Pacific California WoRMS 470514
  229. Smittina palisada
  230. Gordon, 1989 Recent New Zealand WoRMS 470492
  231. Smittina papillifera
  232. (MacGillivray, 1869) Recent SE Australia WoRMS 470485
  233. Smittina? parviovicellosa
  234. Canu & Bassler, 1935 Paleogene Oligocene- Neogene Miocene Australia Victoria.  
  235. Smittina personata
  236. (Hincks, 1884) Recent SE Australia WoRMS 470491
  237. Smittina pileata
  238. (Waters, 1904) Recent Antarctica WoRMS 174383
  239. Smittina pliofistulata
  240. Hayward & Thorpe, 1990 Recent SW Atlantic WoRMS 470509
  241. Smittina pocilla
  242. Hayward & Thorpe, 1990 Recent SW Atlantic WoRMS 174445
  243. Smittina porelloides
  244. (Canu & Bassler, 1920) Paleogene Eocene Priabonian USA, Hungary  
  245. Smittina porosa
  246. Canu & Bassler, 1928 Recent East Atlantic Morocco WoRMS 1043830
  247. Smittina porrigens
  248. (Reuss, 1865) Paleogene Oligocene Rupelian Germany  
  249. Smittina portiuscula
  250. Hayward & Thorpe, 1990 Recent SW Atlantic WoRMS 470510
  251. Smittina pouyetae
  252. el Hajjaji, 1992 Neogene Miocene Messinian Morocco  
  253. Smittina protrusa
  254. Powell, 1967 Recent, Pleistocene Australia WoRMS 470529
  255. Smittina pseudoacutirostris
  256. Gostilovskaya, 1957 Recent Arctic WoRMS 423758
  257. Smittina? pseudocompressa
  258. Androsova, 1958 Recent NW Pacific Sakhalin WoRMS 470534
  259. Smittina? pulchra
  260. Androsova, 1958 Recent NW Pacific WoRMS 470532
  261. Smittina? punctifera
  262. (Canu, 1908) Neogene Miocene Argentina  
  263. Smittina puncturata
  264. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA North Carolina  
  265. Smittina? pupa
  266. Canu & Bassler, 1920 Paleogene Oligocene Rupelian USA Alabama  
  267. Smittina purpurea
  268. (Hincks, 1881) Recent Australia WoRMS 470490
  269. Smittina remotorostrata
  270. (Canu & Bassler, 1928) Recent
    Neogene Miocene Messinian
    Mediterranean, NE Atlantic
    Morocco
    WoRMS 111562
  271. Smittina reptans
  272. (Waters, 1904) Recent Antarctic WoRMS 853629
  273. Smittina reticuloides
  274. Canu & Bassler, 1920 Paleogene Oligocene Rupelian USA Alabama  
  275. Smittina retifrons
  276. Osburn, 1952 Recent N Pacific Alaska WoRMS 470538
  277. Smittina rogickae
    (Homonym)
  278. Hayward & Taylor, 1984 Recent
    Neogene Pleistocene
    Antarctica WoRMS 174386
  279. Smittina rosacea
  280. Powell, 1967 Recent New Zealand WoRMS 470487
  281. Smittina rosefieldensis
  282. McGuirt, 1941 Paleogene Oligocene Rupelian USA Louisiana  
  283. Smittina salsa
  284. Pizzaferri, 2004 Neogene Pliocene-Pleistocene Italy  
  285. Smittina scrupea
  286. Hayward & Taylor, 1984 Neogene Pleistocene Antarctica WoRMS 470524
  287. Smittina seguenzai
  288. (Reuss, 1869) Paleogene Eocene Priabonian Italy  
  289. Smittina sextapuncta
  290. (Canu, 1912) Paleogene Eocene Lutetian France  
  291. Smittina sigillata
  292. (Jullien, 1888) Recent South Atlantic Falklands WoRMS 853911
  293. Smittina sitella
  294. Hayward & Cook, 1983 Recent S Africa WoRMS 470495
  295. Smittina smitti
  296. (Kirchenpauer, 1874) Recent Arctic WoRMS 111564
  297. Smittina smittiana
  298. (Busk, 1884) Recent S Atlantic WoRMS 470511
  299. Smittina smittiella
  300. Osburn, 1947 Recent West Atlantic - Caribbean WoRMS 422353
  301. Smittina sordida
  302. Canu & Bassler, 1920 Paleogene Eocene Priabonian USA Georgia  
  303. Smittina spathulifera
  304. (Hincks, 1884) Recent NE Pacific Canada WoRMS 470554
  305. Smittina spiraminifera
  306. Gordon, 1984 Recent SW Pacific Kermadec WoRMS 470488
  307. Smittina stigmatophora
  308. (Busk, 1884) Recent SW Atlantic WoRMS 470512
  309. Smittina stricta
  310. Vigneaux, 1949 Neogene Miocene Tortonian? France  
  311. Smittina subcordata
  312. Gontar, 1993 Recent NW Pacific WoRMS 470517
  313. Smittina terraenovae
  314. Brown, 1954 Neogene Pliocene
    Recent
    New Zealand WoRMS 470537
  315. Smittina thompsoni
  316. Kluge, 1955 Recent Arctic WoRMS 465715
  317. Smittina torques
  318. Powell, 1967 Recent New Zealand WoRMS 470489
  319. Smittina triangularis
  320. Duvergier, 1921 Neogene Miocene Burdigalian France  
  321. Smittina tuberosa
  322. Kluge, 1952 Recent N Pacific WoRMS 470525
  323. Smittina tubulata
  324. (Gabb & Horn, 1862) Paleogene Oligocene Rupelian USA Alabama  
  325. Smittina umbilicata
  326. (Römer, 1863) Oligocene Miocene Europe  
  327. Smittina undulimargo
  328. Moyano, 1983 Recent Chile WoRMS 470522
  329. Smittina unicus
  330. Tilbrook, 2006 Recent Solomon Islands WoRMS 470557
  331. Smittina uruguayensis
  332. d'Hondt, 1981 Recent SW Atlantic WoRMS 470501
  333. Smittina? uschakowi
  334. Androsova, 1958 Recent NW Pacific WoRMS 470533
  335. Smittina uttleyi
  336. Brown, 1958 Neogene Miocene SE Australia  
  337. Smittina vaciva
  338. (Jullien, 1882) Recent Northeast Atlantic France Bay of Biscay old WoRMS 1043831
  339. Smittina variavicularis
  340. Mawatari, 1974 Recent Japan WoRMS 470527
  341. Smittina veleroa
  342. Soule, Soule & Chaney, 1995 Recent NE Pacific California WoRMS 470513
  343. Smittina volcanica
  344. Moyano, 1983 Recent Chile WoRMS 470523

The list below is of species identified as "Smittia" which appear to have no subsequent revision. They may belong to Smittina, but equally may be species of Parasmittina or Smittoidea.

  1. "Smittia" gemmata
  2. Jullien, 1903 Recent Northeast Atlantic France Gulf of Gascony
  3. "Smittia" grimaldii
  4. Jullien, 1903 Recent Northeast Atlantic France Gulf of Gascony
  5. "Smittia" immersa
  6. Jullien, 1903 Recent Northeast Atlantic France Gulf of Gascony
  7. "Smittia" incisa
  8. Canu, 1904 non (Reuss, 1873) Neogene Argentina
  9. "Smittia" levinseni
  10. Nordgaard, 1903 Recent Arctic
  11. "Smittia" miniacea
  12. Jullien, 1882 Recent Northeast Atlantic France Bay of Biscay
  13. "Smittia" reticulata nitida
  14. MacGillivray, 1895 Neogene Miocene Langhian Australia Victoria
  15. "Smittia" robusta
  16. Jullien, 1903 Recent Northwest Atlantic Grand Banks
  17. "Smittia" schlumbergeri
  18. Jullien, 1903 Recent Northwest Atlantic Grand Banks
  19. "Smittia" subtorquata
  20. Canu, 1904 Neogene Argentina
  21. "Smittia" zuccari
  22. Neviani, 1895 Neogene Pliocene Italy
Smittina kukuiula Smittina nitidissima Smittina papillifera Smittina sp.1
Smittina kukuiula
(Thanks to Dr D. Soule)
Smittina nitidissima Smittina papillifera Smittina sp.1

This information is based upon various sources, but little personal knowledge. I am indebted to the late Alan Horowitz for his compilation, and to Dr. J. Pachut.
I would be grateful for corrections to, or critical reviews of this information. This is not a 'final' version (an impossible ideal), but a working document.

Harmer notebook - entry for Smittia
Harmer notebook - entry for Smittina

Smittina page at NMITA

Smittina bella at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina beringia at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina landsborovii at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina minuscula at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina mucronata at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina pseudoacutirostris at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina smitti at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina thompsoni at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittina tuberosa at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittoidea glaciata at Atlas of Arctic Bryozoa - Piotr Kuklinski.
Smittoidea reticulata at Atlas of Arctic Bryozoa - Piotr Kuklinski.

Smittina hanaishiensis at Pleistocene Bryozoa of Kuromatsunai, Japan.


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Edited by Phil Bock
Modified on 31/3/2017
Modified on 11/11/2017 (Five species added, also "Smittia" spp.)
Modified on 24/1/2018 (Added S. aciculata)
Modified on 24/1/2018 (Removed S. decipiens, S. fallax)
Modified on 12/3/2019 (Moved S. maleposita to Smittoidea)
This URL is http://bryozoa.net/cheilostomata/smittinidae/smittina.html