Escharella Gray, 1848

Type species: Lepralia immersa Fleming, 1828

Synonyms Author Type Species
Haywardipora Soule, Soule & Chaney, 1995 Haywardipora rugosa Soule, Soule & Chaney, 1995
Mucronella Hincks, 1880 Lepralia peachi Johnston, 1847 (=Lepralia immersa)
Perigastrella Hincks, 1880 Discopora coccinea labiata Boeck in Smitt, 1868

This genus includes the following species. (This list is not definitive; many species originally assigned to Mucronella and Perigastrella are included here, but their identity has not been checked. Critical reviews of data are welcome.)

  1. Escharella abyssicola
  2. (Norman, 1869) Recent NE Atlantic WoRMS 111480
  3. Escharella acuta
  4. Zabala, Maluquer & Harmelin, 1993 Recent Mediterranean WoRMS 111481
  5. Escharella anatirostris
  6. (O'Donoghue, 1924) Recent S. Africa WoRMS 470311
  7. Escharella anatirostris spinifera
  8. (O'Donoghue, 1924) Recent S. Africa WoRMS 472711
  9. Escharella areolata
  10. ?      
  11. Escharella areolata parva
  12. (Cipolla, 1928) Tertiary Libya  
  13. Escharella ashapuraae
  14. Guha & Gopikrishna, 2007 Neogene Miocene Aquitanian India Gujarat  
  15. Escharella capitata
  16. (Canu & Lecointre, 1930) Tertiary W. France  
  17. Escharella connectens
  18. (Ridley, 1881) Recent S. Atlantic WoRMS 111482
  19. Escharella? cothurnica
  20. (Kirkpatrick, 1888) Recent Indian Ocean - Mauritius WoRMS 471049
  21. Escharella crozetensis
  22. (Waters, 1904) Recent S. Indian O. WoRMS 470308
  23. Escharella cryptooecium
  24. Souto, Fernandez-Pulpeiro & Reverter-Gil, 2007 Recent East Atlantic WoRMS 470318
  25. Escharella cycloris
  26. (Gabb & Horn, 1862) Paleogene Oligocene USA Alabama  
  27. Escharella diaphana
  28. (MacGillivray, 1879) Recent Australia WoRMS 470305
  29. Escharella dijmphnae
  30. (Kluge, 1929) Recent Arctic WoRMS 470310
  31. Escharella discors
  32. Hayward & Cook, 1983 Recent S. Africa WoRMS 470306
  33. Escharella? elongata
  34. (Canu & Bassler, 1935) Paleogene Oligocene Australia Victoria  
  35. Escharella fistula
  36. Brown, 1952 Tertiary New Zealand  
  37. Escharella fusca
  38. (O'Donoghue & O'Donoghue, 1926) Recent Canada WoRMS 470316
  39. Escharella gilsoni
  40. De Blauwe, 2006 Recent Belgium WoRMS 247907
  41. Escharella granulosa
  42. (Canu & Bassler, 1920) Paleogene Eocene Priabonian USA Mississippi  
  43. Escharella grossa
  44. Moissette, 1988 U. Miocene Algeria  
  45. Escharella grotriani
  46. (Stoliczka, 1862) Paleogene Oligocene Germany  
  47. Escharella guernei
  48. (Jullien & Calvet, 1903) Recent East Atlantic WoRMS 865053
  49. Escharella? hoernesi
  50. (Reuss, 1864) Paleogene Oligocene Germany  
  51. Escharella? hoernesi paroiporosa
  52. Canu & Bassler, 1929 Paleogene Eocene Lutetian Belgium  
  53. Escharella hozawai
  54. (Okada, 1929) Recent Japan WoRMS 470314
  55. Escharella immersa
  56. (Fleming, 1828) Recent North Atlantic, Arctic WoRMS 111484
  57. Mucronella inconspicua
  58. Ortmann, 1890 Recent Northwest Pacific - Japan WoRMS 471051
  59. Escharella incudifera
  60. Gordon, 1984 Recent Kermadec Ridge WoRMS 470302
  61. Escharella indivisa
  62. Levinsen, 1916 Recent Arctic WoRMS 111485
  63. Escharella klugei
  64. Hayward, 1979 Recent N. Atlantic, Arctic WoRMS 111486
  65. Escharella labiata
  66. (Boeck in Smitt, 1868) Recent Atlantic WoRMS 111487
  67. Escharella labiosa
  68. (Busk, 1856) Recent Britain (S) WoRMS 111488
  69. Escharella lagaaiji
  70. Buge, 1957 Tertiary W. France  
  71. Escharella laqueata
  72. (Norman, 1864) Recent NE Atlantic WoRMS 111489
  73. Escharella latodonta
  74. Kluge, 1962 Recent Arctic WoRMS 423358
  75. Escharella levinseni
  76. Hayward, 1994 Recent Faroe Is. WoRMS 111490
  77. Mucronella longicella
  78. Canu, 1914 Paleogene Eocene Lutetian France  
  79. Escharella longicollis
  80. (Jullien, 1882) Recent NE Atlantic WoRMS 111491
  81. Escharella lopezfei
  82. Souto, Berning & Ostrovsky, 2016 Recent Northeast Atlantic Spain WoRMS 868424
  83. Escharella macrodonta
  84. Levinsen, 1917 Recent Arctic WoRMS 990200
  85. Escharella mamillata
  86. Hayward & Thorpe, 1989 Recent Clarence Is. WoRMS 173661
  87. Escharella modica
  88. (Canu, 1926) Paleogene Eocene Bartonian Belgium  
  89. Escharella nilotica
  90. (Ziko, 1985) Eocene Lutetian-Bartonian Egypt  
  91. Mucronella obesa
  92. Faura & Canu, 1916 Paleogene Eocene Priabonian Spain  
  93. Escharella obscura
  94. Norman, 1909 Recent East Atlantic Madeira WoRMS 470317
  95. Escharella octodentata
  96. (Hincks, 1880) Recent NE Atlantic, Mediterranean WoRMS 111492
  97. Escharella ovoidea
  98. (Reuss, 1848) Neogene Miocene Tortonian Austria  
  99. Escharella patens
  100. (Canu & Bassler, 1920) Paleogene Eocene Priabonian USA North Carolina  
  101. Escharella praealta
  102. (Calvet, 1907) Recent W. Europe WoRMS 111493
  103. Escharella quadrata
  104. lopez de la Cuadra & Garcia-Gomez, 2001 Recent Mediterranean WoRMS 221556
  105. Escharella reussiana
  106. (Busk, 1859) Tertiary W. Europe  
  107. Escharella rogeri
  108. David, Mongereau & Pouyet, 1972 Neogene Miocene Burdigalian S. France  
  109. Escharella rugosa
  110. (Soule, Soule, & Chaney, 1995) Recent California WoRMS 470307
  111. Escharella rylandi
  112. Geraci, 1974 Recent Mediterranean WoRMS 111494
  113. Escharella selseyensis
  114. Cheetham, 1966 Eocene England  
  115. Escharella serratilabris
  116. (O'Donoghue, 1924) Recent S. Africa WoRMS 470312
  117. Escharella spinosissima
  118. (Hincks, 1881) Recent SE Australia, New Zealand WoRMS 470304
  119. Escharella takatukii
  120. (Okada, 1929) Recent Japan WoRMS 470315
  121. Escharella tenera
  122. (Reuss, 1874) Eocene Austria, Hungary  
  123. Escharella teres
  124. (Hincks, 1881) Recent Australia WoRMS 470303
  125. Escharella thompsoni
  126. Kluge, 1914 Recent Antarctic WoRMS 465716
  127. Escharella trispinosa
  128. Li, 1990 L. Miocene France  
  129. Escharella? uncifera
  130. (Canu & Bassler, 1929) Recent Philippines WoRMS 470313
  131. Escharella variolosa
  132. (Johnston, 1838) Recent NW Europe WoRMS 111495
  133. Escharella ventricosa
  134. (Hassall, 1842) Recent NE Atlantic, W Mediterranean WoRMS 111496
  135. Escharella ventricosa parva
  136. Buge, 1957 Tertiary W.France  
  137. Escharella vigneauxi
  138. David, 1965 Tertiary France  
  139. Escharella watersi
  140. Hayward & Thorpe, 1989 Recent Antarctica WoRMS 173663

The following species have been assigned to Mucronella, and are included here with the hope that they will be revised eventually.

  1. Mucronella beneckei
  2. Koschinsky, 1885 Paleogene Eocene Lutetian Germany  
  3. Mucronella cretacea
  4. Canu, 1922 Cretaceous Madagascar  
  5. Mucronella formidabilis
  6. Thornely, 1907 Recent Indian Ocean WoRMS 990416
  7. Mucronella cystioides
  8. Koschinsky, 1885 Paleogene Eocene Lutetian Germany  
  9. Mucronella hians
  10. (Hennig, 1892) Cretaceous Sweden  

The following species have been assigned to Perigastrella, and are included here with the hope that they will be revised eventually.

  1. Perigastrella ansata
  2. Canu & Bassler, 1920 Tertiary USA
  3. Perigastrella? costifera
  4. Canu & Bassler, 1920 Tertiary USA
  5. Perigastrella crassilabiata (Homonym?)
  6. Canu & Lecointre, 1930
    Not Duvergier, 1921
    Tertiary W. France
  7. Perigastrella depressa
  8. Canu & Bassler, 1920 Tertiary USA
  9. Perigastrella elegans
  10. Canu & Bassler, 1920 Tertiary USA
  11. Perigastrella exserta
  12. (Gabb & Horn, 1862) Tertiary USA
  13. Perigastrella granulata
  14. Zagorsek, 1994 Eocene Austria, Hungary
  15. Perigastrella helvetica
  16. Canu & Lecointre, 1930 Tertiary W. France
  17. Perigastrella hexagonalis
  18. Canu & Bassler, 1920 Tertiary USA
  19. Perigastrella labiatula
  20. Canu & Lecointre, 1930 Tertiary France
  21. Perigastrella lunata
  22. Canu & Bassler, 1924 Miocene Hungary
  23. Perigastrella lyrulata
  24. Canu & Lecointre, 1930 Tertiary France
  25. Perigastrella maxilla
  26. Canu & Bassler, 1920 Tertiary USA
  27. Perigastrella oscitans
  28. Canu & Bassler, 1920 Tertiary USA
  29. Perigastrella ovalis
  30. Canu & Bassler, 1929 Recent Philippines
  31. Perigastrella ovoidea
  32. Canu & Bassler, 1917 Tertiary USA
  33. Perigastrella? parvicella
  34. Canu & Bassler, 1924 Miocene Hungary
  35. Perigastrella plana
  36. Canu & Bassler, 1920 Tertiary USA
  37. Perigastrella rectilineata
  38. Canu & Bassler, 1920 Tertiary USA
  39. Perigastrella rhomboidalis
  40. Canu & Bassler, 1920 Tertiary USA
  41. Perigastrella serrulata
  42. (Reuss, 1848) Tertiary Europe
  43. Perigastrella sinedorso
  44. Duvergier, 1921 Miocene France
  45. Perigastrella trapezoidea
  46. Canu & Bassler, 1920 Tertiary USA
  47. Perigastrella? tubulosa
  48. Canu & Bassler, 1920 Tertiary USA
Escharella diaphana Escharella ventricosa
Escharella diaphana Escharella ventricosa

Escharella page at NMITA

Escharella immersa at Atlas of Arctic Bryozoa - Piotr Kuklinski.

Escharella macrodonta at Atlas of Arctic Bryozoa - Piotr Kuklinski.

Harmer notebook - entry for Escharella
Harmer notebook - entry for Mucronella
Harmer notebook - entry for Perigastrella


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Edited by Phil Bock
Modified on 25/01/2016
Modified on 20/4/2017
Modified on 17/9/2017
This URL is http://bryozoa.net/cheilostomata/romancheinidae/escharella.html